North Carolina Botanical Garden

Plant Index

Scientific Name:
Common Name:
Genus:
Family:

Dryopteris carthusiana

Dryopteris carthusiana (Vill.) H.P. Fuchs

spinulose wood-fern, toothed wood-fern

Synonym(s): Dryopteris austriaca var. spinulosa, Dryopteris spinulosa

Dryopteris carthusiana (Spinulose Wood-fern)

Image ID: 14192
Image by: Sorrie, Bruce A.
Image Collection: NCBG Digital Library

PLANT INDEX

ID_PLANT: DRCA11
SCIENTIFIC NAME: Dryopteris carthusiana
Include in WOTAS: 0
Publish to Web: 1
Last Modified: 2020-01-01

GENUS INDEX

GENUS CODE: DRYOP
GENUS SCIENTIFIC: Dryopteris
GENUS AUTHORITY: Adans.
GENUS COMMON: Woodfern
GENUS SUMMARY: A genus of about 400 species, nearly cosmopolitan, but concentrated in temperate Asia. Sessa, Zimmer, & Givnish (2012) discuss the phylogeny and biogeography of Dryopteris; the clades shown in the key are from their work.
GENUS IDENTIFICATION: Identification notes: Dryopteris and Athyrium are often confused when not fertile; they can be easily distinguished by breaking off a leaf and counting vascular bundles (which will appear as thread-like strands). Dryopteris has 5 and Athyrium has 2. Many Dryopteris species will hybridize with one another to form sterile hybrids. Whenever two or more Dryopteris species are found growing together, there is a good chance that hybrids are present. Hybrids generally show intermediacy between the two parents, and have abortive sporangia or spores.
GENUS REFERENCES: Montgomery & Wagner in FNA (1993b); Sessa, Zimmer, & Givnish (2012); Montgomery & Paulton (1981); Montgomery (1982); Kramer et al. in Kramer & Green (1990); Hoshizaki & Wilson (1999).

FAMILY INDEX

FAMILY CODE: DRYOPT
FAMILY SCIENTIFIC: Dryopteridaceae
FAMILY AUTHORITY: Ching 1965
FAMILY COMMON: Wood-fern Family
FAMILY SUMMARY: A family of about 40-45 genera and 1700 species, cosmopolitan in distribution, but concentrated in temperate and montane areas. Here circumscribed (following Smith et al. 2006) to exclude Onocleaceae and Woodsiaceae.
FAMILY REFERENCE: Smith in FNA (1993b); Smith et al. (2006); Lellinger (1985); Kramer et al. in Kramer & Green (1990).

NCBG DESCRIPTIVES

INTRO:
STEMS:
LEAVES:
INFLORESCENCE:
FLOWERS:
FRUITS:
COMMENTS:
HEIGHT:

DURATION: Perennial
HABIT: Herb, Fern

LEAF ARRANGEMENT:
LEAF COMPLEXITY:
LEAF RETENTION: Deciduous

FLORAL CHARACTERISTICS
SYMMETRY:
BLOOM TIME: Jun-Sep

Jan	Feb	Mar	Apr	May	Jun	Jul	Aug	Sep	Oct	Nov	Dec
					x	x	x	x

BLOOM COLOR:

White	Red	Pink	Orange	Yellow	Green	Blue	Lavender	Purple	Violet	Brown	Not Applicable
											x

FRUITING PERIOD:

DISTRIBUTION
HABITAT TYPE:
NATIVE RANGE:

HORTICULTURAL
Plant Sale Text:

Bloom Table Text:

NCBG Location:

Cultural Notes:

SOIL MOISTURE: Average, Moist/Wet
LIGHT EXPOSURE: Sun, Part Shade, Shade
MINIMUM HARDINESS ZONE:
MAXIMUM HARDINESS ZONE:
GERMINATION CODE:
WILDLIFE VALUE:
DEER RESISTANCE:

USDA PLANTS DATABASE

USDA Symbol: DRCA11
USDA Common Name: Spinulose Woodfern
Native Status: L48 (N), CAN (N), SPM (N)
Distribution: USA (AL, AR, CT, DC, DE, GA, IA, ID, IL, IN, KY, MA, MD, ME, MI, MN, MO, MT, NC, ND, NE, NH, NJ, NY, OH, OR, PA, RI, SC, SD, TN, VA, VT, WA, WI, WV), CAN (AB, BC, LB, MB, NB, NF, NS, NT, NU, ON, PE, QC, SK, YT), FRA (SPM)
Duration: Perennial
Growth Habit: Forb/herb

NATIONAL WETLAND INDICATOR STATUS

Region:	AGCP	AK	AW	CB	EMP	GP	HI	MW	NCNE	WMVE
Status:	FACW		FAC		FAC	FACW		FACW	FACW	FAC

This information is derived from the U.S. Army Corps of Engineers 2016 National Wetland Plant List, Version 3.3 (Lichvar, R.W., D.L. Banks, W.N. Kirchner, and N.C. Melvin. 2016. The National Wetland Plant List: 2016 wetland ratings. Phytoneuron 2016-30: 1-17. Published 28 April 2016. ISSN 2153 733X). Regions: AGCP-Atlantic and Gulf Coastal Plain, AK-Alaska, AW-Arid West, CB-Caribbean, EMP-Eastern Mountains and Piedmont, GP-Great Plains, HI-Hawaii, MW-Midwest, NCNE-Northcentral and Northeast, WMCV-Western Mountains, Valleys & Coast

2018 Natural Heritage Program List of Rare Plant Species of North Carolina

NHP ID	S RANK	ROUNDED S RANK	G RANK	ROUNDED G RANK	STATE STATUS	FEDERAL STATUS
16291	S2	S2	G5	G5	W7

This information is derived from the 2018 North Carolina Natural Heritage Program List of Rare Plant Species.

WEAKLEY FLORA

SCIENTIFIC NAME: Dryopteris carthusiana
COMMON NAME: Spinulose Wood-fern, Toothed Wood-fern
SYNONYMY: [= Ar, C, FNA, Il, K, Pa, Tn, Va, W; = D. spinulosa (O.F. Mueller) Watt - RAB, S, WV; = D. spinulosa var. spinulosa - F; = D. austriaca (Jacquin) Woynar ex Schinz & Thellung var. spinulosa (O.F. Mueller) Fiori - G]
PHENOLOGY: Jun-Sep.
HABITAT: Acidic, organic-rich bogs, swamps, less frequently in moist rocky ravines, rich forests, and sloping rock outcrops.
COMMENTS: Irregularly circumboreal, in North America ranging from n. QC west to YT, south to NC, SC, ne. GA, TN, AR, NE, w. MT, and WA. This species is a fertile allotetraploid derived from hybridization of D. intermedia and D. semicristata, a hypothetical species which may now be extinct. Its chromosome complement is symbolized IISS.
RANGE MAP: Dryopteris carthusiana.png

Key to Map Symbols

ABOUT FAMILY (Weakley Flora)
Dryopteridaceae Ching 1965 (Wood-fern Family)
SUMMARY: A family of about 40-45 genera and 1700 species, cosmopolitan in distribution, but concentrated in temperate and montane areas. Here circumscribed (following Smith et al. 2006) to exclude Onocleaceae and Woodsiaceae.
REFERENCE: Smith in FNA (1993b); Smith et al. (2006); Lellinger (1985); Kramer et al. in Kramer & Green (1990).

ABOUT GENUS (Weakley Flora)
Dryopteris Adans. (Woodfern)
SUMMARY: A genus of about 400 species, nearly cosmopolitan, but concentrated in temperate Asia. Sessa, Zimmer, & Givnish (2012) discuss the phylogeny and biogeography of Dryopteris; the clades shown in the key are from their work.
REFERENCE: Montgomery & Wagner in FNA (1993b); Sessa, Zimmer, & Givnish (2012); Montgomery & Paulton (1981); Montgomery (1982); Kramer et al. in Kramer & Green (1990); Hoshizaki & Wilson (1999).

HERBARIUM RESOURCES

SERNEC: Find Dryopteris carthusiana in Southeast Regional Network of Experts and Collections (if available)
UNC SERNEC: Find Dryopteris carthusiana in University of North Carolina Chapel Hill Herbarium - Southeast Regional Network of Experts and Collections (if available)

WEB RESOURCES

USDA: Find Dryopteris carthusiana in USDA Plants
NPIN: Find Dryopteris carthusiana in NPIN Database
FNA: Find Dryopteris carthusiana in the Flora of North America (if available)
Google: Search Google for Dryopteris carthusiana

IMAGE USE RECORDS

ID IMAGE: 14192 - Primary Image FloraQuest Plant Detail Page (Landscape Preferred)
ID IMAGE1: 14192 - Primary Image WOTAS
ID IMAGE2: 14192 - Secondary Image WOTAS
ID IMAGE3: 0 - Primary Image Plant Sale Sign (Landscape Only)
ID IMAGE4: 0 - Primary Image New Plant Sale Label (Portrait Only)